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Settlement abundances of barnacles are determined by larval supply and the ratio of settlement to larval supply (K) This ratio can be divlded into 2 funchons the ratlo of settlement on bare space to larval supply (k , ) , and the loss of cypnds due to the preempting of rock space by sessde orgamsms (k,) To exarmne the 3 functions of settlement ( l e larval supply, k , and k, ) of the barnacle Sembalanus Carlosus, we monitored the daily water column larval supply and the dally settlement density in plots where all benthos were removed and in natural-community plots, which were leit undisturbed, dunng one settling season In both lowand mid-tidal sites, k , explained >?5% of day-to-day variabdity in K k , and k2 were density dependent, whde daily settlement densities were linearly related to larval supply k , varied with tidal height and dady larval supply, but not wlth other physical vanables k2 vaned with daily settlement densities in community-removal plots and was not related to any physical or biolog~cal vanable Such dens~ ty dependence in k, and k2 may have been caused by 3 processes (1) inhibition of larval settlement by planktonic larvae, (2) enhancement of settlement by conspec~flcs (and t h e ~ r chenlical attractants) on days wlth relatively low larval density, and/or (3) preemptlon of suitable settlement substrata by settlers that attached on days w ~ t h relatively high larval density

Daily settlement variability of the barnacle Semibalanus cariosus:importance of physical factors and density-dependent processes