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Echinogamrnarus pirloh and E. obtusatus are 2 species of amphipod that inhablt boulder shores. Their distribution on such shores on Great Cumbrae Island, Scotland, was found to be zoned; E. obtusatus always occurring from MTL downwards and E. pirloti living between MTL and HWN (summer) or HWS (winter). Prevlous investigation of this shore established that oxygen tension, temperature, salinity and redox potential were the most variable environmental parameters of the amphipods' understone habitat and that there were cons~derable differences between upper and lower shore sites. E. pirloti was more tolerant of anoxia, high temperature and low salinity than E. obtusatus. The levels of oxygen tension and temperature that induced physiological 'stress' in the laboratory were similar to those recorded on the shore, but both species were able to survive indefinitely in the lowest salinities recorded on the beach. There was no difference in tolerance of low humidity between the 2 species, and there was similarly no difference between understone hurnidities recorded on the upper and lower shore. The 'dominant' factors determining zonation on the shore are probably tolerance of oxygen tension and high temperature.

Effects of oxygen tension, temperature, salinity, and humidity on the survival of two intertidal gammarid amphipods