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Abstract: Examination of previously described and some undescribed leaves of Ulmus and Zelkova from the later Oligocene, Miocene, and Pliocene of western North America indicates that at least eight species of Ulmus and two species of Zelkova are represented. Three new species are described: Ulmus chaneyi, U. knowltoni, and Zelkova browni.

Revisions of Ulmus and Zelkova in the middle and late Tertiary of western North America