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Bl Introduction 1 Stratigraphy of the Puget Group 2 Description of the Field Area 4 Data Collection 5 Stratigraphic Sections 5 Plant-Fossil Collections 6 Adjustments to the Data Set 6 General Characteristicsof the Upper Ravenian Flora 8 Analysis of Plant Distribution Presence-Absence Data 9 Species Richness 10 Species Diversity 10 Leaf Margins of AngiospermTaxa 10 Leaf Size of AngiospermTaxa 12 Conifers, Pteridophytes, and Sphenopsids 12 Comparison of Floristic Assemblages 13 Summary of Data on Presence-Absence of Species 13 Analysis of Plant Data Taxon Abundance  15 Ordination of Localities  15 Methods 15 Results and Discussion 15 Cluster Analysis 16 Methods 16 Results and Discussion 16 Environmental Heterogeneity in the Puget Group 17 Discussion 17 Species Distribution in the Upper Ravenian Puget Group 17 Anomalous Localities and their Use in Interpreting Paleoecology 20 Conclusions 20 Depositional Environments Implications for their Use in Paleobotany 21 Biostratigraphy 21 Paleoclimatology 21 Paleoecology 21 Recommendations for Cenozoic Paleobotanical Investigations : 22 References Cited 22 Appendix 1. Matrix of Abundance Data for 20 Plant-Fossil Localities from the Puget Group 28 Appendix 2. Matrix of Sorenson's Index of Similarity for Comparisons Between 20 Plant-Fossil Localities from the Puget Group 30 Appendix 3. Matrix of Spearman's Rank-Order Correlation Coefficients for 20 Plant-Fossil Localities from the Puget Group 30 Appendix 4. Abbreviations and Taxonomic Citations for Plant-Fossil Taxa from the 20 Localities Examined in the Puget Group 31

Paleoecological and floristic heterogeneity in the plant-fossil record; an analysis based on the Eocene of Washington