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iii GENERAL INTRODUCTION 1 References 5 CHAPTER 1. Persistence and degradation of maize-expressed vaccine protein, Escherichia coli heat-labile enterotoxin subunit B (LTB), in soil and water Abstract 9 Introduction 10 Materials and Methods 13 Results and Discussion 22 Acknowledgements 34 References 34 Tables (1 – 3) 39 Figures (1 – 18) 42 CHAPTER 2. Sub-acute effects of maize-expressed vaccine protein, Escherichia coli heat-labile enterotoxin subunit B (LTB), on the springtail, Folsomia candida, and the earthworm, Eisenia fetida Abstract 60 Introduction 61 Materials and Methods 63 Results and Discussion 70 Acknowledgement 75 References 75 Tables (1 – 2) 79 Figures (1 – 5) 80 GENERAL CONCLUSION 83 References 85

Environmental studies on the fate and effects of pharma proteins produced in transgenic maize