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The C-terminal end of Cdt2 contains a PIP box for binding to PCNA to promote CRL4 Cdt2  function, creating a new paradigm where the substrate receptor and substrates bind to a common multivalent docking platform for ubiquitination.

Direct binding of Cdt2 to PCNA is important for targeting the CRL4Cdt2 E3 ligase activity to Cdt1

Direct binding of Cdt2 to PCNA is important for targeting the CRL4^Cdt2 E3 ligase activity to Cdt1