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4 Introduction 6 Review of Literature 8 Mesenchymal Stem Cells 8 Establishment 8 Differentiation of MSC into Bone, Muscle, and Adipose Tissue 8 Roles in the Adult 9 Poor Maternal Nutrition 10 Effects on Mesenchymal Stem Cells 10 Effects on Bone, Muscle, and Adipose Development 11 Effects on Whole Body Function 13 Fetal Programming and Maternal Diet 13 Metabolism 17 Normal Metabolic Physiology 17 Cell Differentiation Status and Metabolic Preference 18 Regulation of Metabolic Status: Role of Oxygen/Nutrient Sensing Pathways ....21 Reactive Oxidative Species: Physiological and Pathological Levels 23 Altered Metabolic Physiology 25 Conclusion 27 Materials and Methods 29 Results 36

0201 Poor maternal nutrition during gestation alters mesenchymal stem cell (MSC) metabolism in offspring