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The evolutionary history of Ophioglossaceae is enigmatic mainly because fossils of the family trace back only from the earliest Tertiary. Phylogenetic analyses indicate that Helminthostachys is sister to the broadly defined Botrychium. Generally the sporophore of Botrychium is a pinnately compound, whereas it is simple in Helminthostachys. Here examples of different Helminthoistachys are represented which show double or triple spikes with some variations. Plants showing variations in their spike morphology are also grown normally. Variations of Helminthostachys spike morphology indicate a tendency to form a compound sporophore structure and in that way have a strong relationship with Botrychium. Izvleček Evolucijski razvoj družine kačjejezikovk (Ophioglossaceae) je precej skrivnosten, saj prve fosilne ostanke predstavnikov družine najdemo šele iz zgodnjega terciarja. Filogenetske raziskave kažejo, da je monotipični rod Helminthostachys sestrski sicer širše definiranemu rodu mladomesečin (Botrychium). Na splošno je fertilni ali plodni del sporotrofofila pri rodu Botrychium pernato deljen, medtem ko je pri rodu Helminthostachys valjast in enostaven. V študiji so predstavljeni različni osebki vrste Helminthostachys zeylanica z enim ali več plodnimi izrastki. Osebki, ki kažejo na variacijo v obliki trosnih izrastkov rastejo v naravnem okolju. Različna morfologija plodnega-trosnega dela sporotrofofila pri rodu Helminthostachys nakazuje tendenco k tvorbi pernato deljenih trosnih izrastkov, kar je znak sorodnosti omenjenega rodu z rodom Botrychium.

Morphological Varia Tions of Fertile Spike in Helminthostachys Zeylanica (L.) Hook