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Typification of Sydneia manleyi Pšenička et al. 2003 is basedentirely on fertile foliage. Of late, attached sterile-fertile segments havebeen found which are illustrated and described, together with cutic-ularpreparations. A new subfamily, Sydneideae subfam. nov. is erected for themonotypytic species. The synangial chemistry is compared with marattialeansynangia/sporangia from the Czech Republic and from Canada.

Sterile Foliage of Fertile Sydneia Manleyi and Synangial Chemistry (Eusporangiate Fern, Late Asturian, Canada): A New Subfamily Sydneideae