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Age is known to influence fecundity in many avian species. Rockwell et al. (1983) showed that the clutch size of female Lesser Snow Geese (Chen caerulescens caerulescens) increases gradually up to age five, after which there are no detectable age effects. Furthermore, they documented considerable yearly variation in the mean clutch size at the La P6rouse Bay colony near Churchill, Manitoba; this variation could not be attributed to annual differences in the age composition of the breeding females. Instead, they suggested that some temporally variable environmental condition(s) might be affecting the size of the initial clutch laid, the level of partial clutch loss, or some combination of the two. A question that arises is whether the proximate factors resulting in these yearly changes in mean fecundity affect all age classes to the same extent. Are, for example, young and inexperienced females less able to cope with the exigencies of particularly severe breeding seasons? Such a finding would require refinement of the emerging theories that relate average reproductive output of young females to general life-history tactics (e.g., Curio 1983). In addition, a general dependence of yearly fecundity differences on age would complicate models of population dynamics of this species (cf. Findlay et al. 1982). In this paper, we examine this issue by evaluating annual variation in the difference between mean clutch sizes of twoyear-old and older females.

Life History Studies of the Lesser Snow Goose V.: Temporal Effects on Age-Specific Fecundity