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L. amplexicaule L. grows in different regions of Iran and forms several   populations that can be invasive in some area. Morphological and genetic   variation of L. amplexicaule in Iran was studied to evaluate the   differentiation among three presumed varieties within this species.   Morphometric study was accompanied by the analysis of ISSR and chloroplast   genome (psbA-trnH). Results of the molecular methods show that L.   amplexicaule var. bornmuelleri Mennema. does not form any genetically   distinct group. Genetic study of 103 plant specimens of 13 different   populations using ISSR molecular marker revealed a high degree of   within-population (67%) and among (33%) populations genetic diversity. AMOVA   and Gst analyses produced a significant difference. Nm estimation revealed   certain degree of gene flow/shared alleles between these populations,   indicating that L. amplexicaule is predominantly an out-crossing species in   Iran. Mantel test produced significant correlation between genetic distance   and geographical distance of the studied populations (r = 0.097, P = 0.04).   STRUCTURE plot identified two main gene pools for this species in Iran.

Biosystematic study and population genetic analysis in Lamium amplexicaule L. (Lamioideae, Lamiaceae)