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Azhdarchid pterosaurs existed during the Mesozoic era and died out during the mass extinction at the end of the Cretaceous. These amazingly large creatures have sparked a debate on whether they were capable of flying or simply gliding due to their massive size, anatomy, and morphology. Two theories that have been developed are that these creatures were terrestrial stalkers and that they are closer to pelicans in terms of feeding capability. More specifically, Naish and Witton suggest that these giants were capable of covering large distances by soaring and could forage on land by walking quadrupedally using their long necks to reach down and kill small animals, consume dead tissues and possibly fruits. Fossil evidence to support this theory includes tracks of the Azhdarchid’s showing a parasagittal gait, a winged planform, and specimens have been only found in continental depositional settings. Evidence also indicates that these creatures had membranes serving as wings, but it cannot be discerned with current knowledge whether membranes were used for gliding or for true flying.

A Summary of Current Research on the Functional Morphology of Flight in Azhdarchid Pterosaurs