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Two new western Atlantic species of Gobionellus (family Gobiidae) are described and figured. G. comma, which is presently known only from the southern Caribbean Sea, off Venezuela, is characterized primarily by a dark, comma-shaped bar in the suborbital area. G. atripinnis, which has been found only in the western Gulf of Mexico, from southern Texas to· Veracruz, Mexico, is most readily distinguished by an elongate black blotch in the male's spinous dorsal fin and in having 16 pectoral fin rays. G. comma is closely related to the eastern Pacific G. manglicola The relationships of G. atripinnis are more obscure. Important diagnostic characters of Gobionellus are presented, together with preliminary conclusions concerning the interrelationships of the genera Evorthodus and Oxyurichthys, which closely resemble Gobionellus in several important ways. Although groundwork is laid for possible synonymization of these genera (including discussion of nomenclatural problems), such action is deferred until more comprehensive studies are completed. A total of 14 coarse-scaled species of Gobionellus are recognized. These are included in a taxonomic key, which also includes a geographic range statement for each species. Also included in the key are the two species of Evorthodus, which are frequently confused with the coarse-scaled Gobionellus. This key does not include the recently-described G. munizi Vergara 1978, specimens of which we have not had the opportunity to examine. Comments are included, however, regarding this species' validity and probable relationships, based on text of the original description and accompanying figures, The genus Gobionellus Girard was reviewed by Ginsburg (1932), who recognized a total of 11 species in the western Atlantic and eastern Pacific oceans, plus two (in the subgenus Biat) in the IndoPacific region. Several other new species were subsequently described (Pfaff, 1933; Ginsburg, 1953; Mead and Bohlke, 1958; Gilbert and Randall, 1968; and Gilbert and Randall, in Gilbert and Kelso, 1971), and these, together with several previously described forms later recognized by Ginsburg (1953), increased the total to 20. One recently named species (Gobionellus pseudofasciatus Gilbert and Randall), for which only a brief diagnosis was originally provided (Gilbert and Kelso, 19 71), has 27 since been described in greater detail (Hastings, 1979) particularly with regard to color and pigmentary variation. We have been engaged for some time in a review of the "coarse-scaled" species of Gobionellus (i.e., those forms having 46 or fewer scales in the lateral series). Although final publication of this work is not yet ready, we consider it desirable to accord formal status to the two remaining undescribed coarse-scaled species, and to provide a key (with range statements) to the recognized species. We also present evidence suggesting a close relationship to Gobionellus of the genera Evorthodus and Oxyurichthys, but stop short of synonymization, primarily on the recommendation of 1 Gilbert and Randall: Two New Western Atlantic Species of the Gobiid Fish Genus Gobione Published by The Aquila Digital Community, 1979 28 C. R. Gilbert and J. E. Randall Douglass F. Roese, of the Australian Museum. Dr. Roese, who is actively working on the systematics of IndoPacific gobies, agrees that these groups likely are closely related, but points out (in lift.) that basic morphological differences exist among Oxyurichthys, coarse-scaled Gobionellus, fine-scaled Gobionellus and Evorthodus. He also notes that complexities remain concerning relationships of these groups to certain other genera (e.g., Waitea, Oligolepis and Paroxyurichthys). Inasmuch as further study is necessary before these relationships can be completely resolved, Dr. Roese advocates a conservative approach regarding nomenclatural changes. Although this leaves the status of certain species (notably Gobionellus stigmalophius) temporarily in limbo, we feel that this is preferable to making changes that might ultimately

Two New Western Atlantic Species of the Gobiid Fish Genus Gobionellus, with Remarks on Characteristics of the Genus