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Since it was found that M and N are the same important signs of blood as the 4 classical groups (A, B, AB and O), since they are inherited according to Mendel's law, the question had to arise about using them to establish paternity.

Olbrich S. On the use of adsorbed immune sera for the diagnosis of M- and N-blood groups. (Ztschr. F. Jmmunitätsf., Bd. 90, 1937)