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1School of Biological, Earth and Environmental Sciences, Distillery Fields, North Mall, University College Cork, Ireland 2SCNU-UCC Joint Laboratory for Ecology and Environmental Science, No.55, Zhongshan Avenue West, Tianhe District, School of Life Science, South China Normal University, Guangzhou 510631, P.R. China Article Information Received 27 December 2019 Revised 21 February 2020 Accepted 13 April 2020 Available online 14 October 2020

Impact of Weather Changes and Human Visitation on the Behavior and Activity Level of Captive Humboldt Penguins