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FUNDING: DSI/NRF Centre of Excellence in Palaeosciences, DSI/NRF African Origins Platform, National Research Foundation (South Africa), the Palaeontological Scientific Trust (PAST), the Millennium Trust Coombs Hill, a new fossil locality in the Witpoort Formation (Witteberg Group) of South Africa, preserves a record of Famennian (Late Devonian) life in Gondwana. Fossil plants collected at Coombs Hill are preliminarily assigned to several classes. Shelly invertebrates include a variety of bivalve mollusc forms, some of which appear to be preserved in life position. Biodiversity at Coombs Hill is comparable to that of the well-known Waterloo Farm lagerstätte in ordinal diversity, but exhibits differences in species composition. Ongoing taxonomic analysis will provide a rare window into the ecology of high-latitude environments during this pivotal stage of Earth history, which immediately preceded the end-Devonian extinction. Sandstone dominated sedimentary facies at Coombs Hill suggest a high-energy coastal marine setting, with brackish back-barrier estuarine/lagoonally derived fossiliferous mudstones. Exact stratigraphic placement within the Witpoort Formation is hampered by structural deformation, and precise age comparisons with Waterloo Farm are currently tenuous.

Coombs Hill: A Late Devonian fossil locality in the Witpoort Formation (Witteberg Group, South Africa)