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The nymph, female imago and egg of Siphluriscus chinensis Ulmer are described and figured in detail for the first time based on associated nymphs and imagos from China. Additional characters of the male imago are described. Based upon imaginal and nymphal characters, Siphluriscidae fam. n. is established in the Siphlonuroidea for Siphluriscus and, provisionally, the Jurassic genus Stackelbergisca Tshernova. Siphluriscus ? davidi Navás, known from one male subimago, is transferred to Siphlonurus davidi comb. n. The family Siphluriscidae is distinguished from all other families of Siphlonuroidea by the presence of coxal gills on the fore and middle legs in all stages and in the nymph by the presence of a basal, moveable, dactyl-like appendage on each claw, and labial and maxillary gills. Characters of the nymphal mouthparts, particularly the mandibles and maxillae, suggest that Siphluriscidae is a sister-group of the family Nesameletidae. However, the majority of character states are considered plesiomorphic in the Ephemeroptera, and may contribute valuable information on the origin, phylogeny and biogeography of Ephemeroptera.

the florida entomologist

THE NYMPH OF SIPHLURISCUS CHINENSIS AND ADDITIONAL IMAGINAL DESCRIPTION: A LIVING MAYFLY WITH JURASSIC ORIGINS (SIPHLURISCIDAE NEW FAMILY: EPHEMEROPTERA)