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L.) chromosomes were first charac-terized by McClintock (1929) from studies of the first pol-len mitosis. Later, McClintock (1930, 1931, 1933) foundthat pachytene chromosomes showed better morphologicaldetails for cytological studies. The pachytene chromosomeswere identified on the basis of their length, centromere po-sition, prominent chromomeres and heterochromatic knobs(Longley, 1938; Rhoades, 1955; McClintock

Heterochromatin of maize chromosomes: structure and genetic effects