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i LIST OF TABLES iv LIST OF FIGURES vi CHAPTER 1: Overview & General Background 1 CHAPTER 2: Woodpecker Abundance & Habitat Use in Urban Parks Abstract 6 Introduction 7 Methods 11 Results 17 Discussion 20 Conclusion 23 CHAPTER 3: Minimum Area Requirements of Woodpeckers in an Urban Landscape Abstract 33 Introduction 34 Methods 38 Results 42 Discussion 44 Conclusion 49 CHAPTER 4: Final Conclusions 64 REFERENCES 66

Woodpeckers in the City: Habitat Use and Minimum Area Requirements of Woodpeckers in Urban Parks and Natural Areas in Portland, Oregon