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Vacuolar type ATPases have been found on various endomem branes of eukaryotic cells, e.g. lysosomes, chromaffin granules, vesicles derived from the Golgi apparatus, endosomes and vacuoles. Although this ATPase type is targeted to different compartments in one cell, only one gene for each subunit had been found per genome.

Two Separate Genes Encode the Catalytic 70 kDa V-ATPase Subunit in Psilotum and Equisetum