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1. Fluorescence changes of umbelliferone have been used as indicator for pH changes in the outer phase of the functional membrane of photosynthesis. In combination with the repetitive pulse technique it is possible to measure the pH change which is produced during one turn-over of the molecular machinery (Δ pH ≈ 10 −4 ). The analysis of this primary pH changes gives insight in the primary ionic events in general.

Primary Ionic Events in the Functional Membrane of Photosynthesis: Umbelliferone as Indicator for pH Changes in one Turn-Over