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In this work, we identified non-crustose invasive ( Codium fragile  subsp.  fragile ) and native  Codium  spp. ( Codium tomentosum  and  Codium vermilara ) in the central Cantabrian Sea using DNA barcoding (tufA and rbcL genes). We designed a new FCO tuf A genetic marker for identifying  Codium  spp. in fresh and herbarium material. The tufA and rbcL sequences revealed three different single haplotypes for each of the species and a lack of intraspecific genetic diversity. The FCO tuf A genetic marker revealed one new haplotype of  C. fragile  within the native region (South Korea), suggesting the possibility of higher genetic diversity in the donor region of this invasive species.

DNA barcoding-based assessment of the invasive and native non-crustose Codium species in the central Cantabrian Sea, southern Bay of Biscay