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Plants of the genus Bulbophyllum Thouars have some of the most intricate flowers and complex pollination mechanisms to be found amongst the Orchidaceae (van der Pijl and Dodson, 1969; van der Cingel, 2001). For an up-to-date, overall account of the genus, readers are referred to Pridgeon et al. (2014). Many of its members are pollinated by dipterans (van der Pijl and Dodson, 1969; van der Cingel, 2001) attracted by foodrewards and fragrance. However, to date, these food-rewards have been investigated for relatively few species, mostly Neotropical and African taxa, and have generally been found to be lipid-rich and produced by palisade-like epidermal cells of the

Evidence for the Dual Role of Floral Secretory Cells in Bulbophyllum