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The placoderm fauna of the upper Famennian tetrapod-bearing locality of Strud, Belgium, includes the antiarch  Grossilepis rikiki , the arthrodire groenlandaspidid  Turrisaspis strudensis  and the phyllolepidid  Phyllolepis undulata . Based on morphological and morphometric evidence, the placoderm specimens from Strud are predominantly recognised as immature specimens and this locality as representing a placoderm nursery. The Strud depositional environment corresponds to a channel in an alluvial plain, and the presence of a nursery in such environment could have provided nutrients and protection to the placoderm offspring. This represents one of the earliest pieces of evidence for this sort of habitat partitioning in vertebrate history, with adults living more distantly from the nursery and using the nursery only to spawn or give live birth.

Placoderm Assemblage from the Tetrapod-Bearing Locality of Strud (Belgium, Upper Famennian) Provides Evidence for a Fish Nursery