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Seven  Scolopendra  species from the Southeast Asian mainland delimited based on standard external morphological characters represent monophyletic groups in phylogenetic trees inferred from concatenated sequences of three gene fragments (cytochrome  c  oxidase subunit 1, 16S rRNA and 28S rRNA) using Maximum likelihood and Bayesian inference. Geometric morphometric description of shape variation in the cephalic plate, forcipular coxosternite, and tergite of the ultimate leg-bearing segment provides additional criteria for distinguishing species. Colouration patterns in some  Scolopendra  species show a high degree of fit to phylogenetic trees at the population level. The most densely sampled species,  Scolopendra dehaani  Brandt, 1840, has three subclades with allopatric distributions in mainland SE Asia. The molecular phylogeny of  S .  pinguis  Pocock, 1891, indicated ontogenetic colour variation among its populations. The taxonomic validation of  S .  dawydoffi  Kronmüller, 2012,  S .  japonica  Koch, 1878, and  S .  dehaani  Brandt, 1840, each a former subspecies of  S .  subspinipes  Leach, 1814 sensu Lewis, 2010, as full species was supported by molecular information and additional morphological data. Species delimitation in these taxonomically challenging animals is facilitated by an integrative approach that draws on both morphology and molecular phylogeny.

plos one

The Centipede Genus Scolopendra in Mainland Southeast Asia: Molecular Phylogenetics, Geometric Morphometrics and External Morphology as Tools for Species Delimitation