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A new species of Bythinella: B. walensae is described from Aria spring at Naxos Island, Greece. Cytochrome oxidase subunit I (COI) sequences of mtDNA, as well as internal transcribed spacer (ITS-1) of nuclear ribosomal DNA indicate distinctness of B. walensae. The shell, female reproductive organs and penis are described. The most characteristic of this species are: slender shell, narrow aperture, J-shaped bursa coupled with bulky and spherical receptaculum, and the length ratio of penis arms. Simple anatomy coupled with wide variation ranges overlapping between the species are emphasised. key words: molecular distinctness, mtDNA, cytochrome oxidase subunit I (COI), ITS-1, morphological distinctness, shell, penis, female reproductive organs

A new species of Bythinella Moquin-Tandon, 1855 (Caenogastropoda: Truncatelloidea) from Naxos Island, Greece