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A MULTIVARIATE MIXED HIDDEN MARKOV MODEL FOR BLUE WHALE BEHAVIOUR AND RESPONSES TO SOUND EXPOSURE1 BY STACY L. DERUITER∗,†, ROLAND LANGROCK‡,†, TOMAS SKIRBUTAS†, JEREMY A. GOLDBOGEN§, JOHN CALAMBOKIDIS¶, ARI S. FRIEDLAENDER‖,∗∗ AND BRANDON L. SOUTHALL∗∗ Calvin College∗, University of St Andrews†, Bielefeld University‡, Stanford University§, Cascadia Research Collective¶, Oregon State University‖ and SEA, Inc.∗∗

A multivariate mixed hidden Markov model for blue whale behaviour and responses to sound exposure