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The dendrobatid genus Silverstoneia is a clade of forest-dwelling frogs that share small adult size (<22 mm snout-vent length), basal webbing between toes III–IV, a solid blackish brown flank divided by a conspicuous pale oblique lateral stripe from the groin to the eye, a pale ventrolateral stripe, finger I > II, male nurse frogs (known in four species), and larvae possessing an umbelliform oral disc with few to many submarginal papillae and medially emarginate posterior labium (also known in four species).\ud\udThe clade is distributed from Costa Rica to southern Departamento del Valle del Cauca in western Colombia and includes eight species, five of which are described herein as new species. As in all species of Dendrobatoidea, in Silverstoneia the distal tendon of insertion of the m. semitendinosus inserts dorsad to the distal tendon of the mm. gracilis complex and is strapped to the dorsal edge of the inner surface of the mm. gracilis complex by a unique binding tendon. The species of Silverstoneia may be diagnosed on the basis of adult size, adult ventral coloration, thigh coloration, and degree of expansion of finger III in adult males; additionally, there are clear species differences among the known tadpoles.\ud\udTaxonomic comments are given for three previously named species: Silverstoneia erasmios (Rivero and Serna), S. flotator (Dunn), and S. nubicola (Dunn). We were unable to distinguish erasmios from nubicola. However, only females of erasmios are known and its validity needs confirmation. The species S. flotator sensu lato and S. nubicola sensu lato occur north through Panama to Costa Rica; they are distinct from one another, but some intraspecific variation suggests the possible presence of unnamed sibling species. The specimen long recognized as the type of S. flotator is not the holotype, which we consider lost; however, ample material is available from the well-known type locality (Barro Colorado Island) and neotype designation is not needed.\ud\udFive new species are described—all endemic to low and moderate elevations on the Pacific versant of Colombia: Silverstoneia dalyi (p. 5), S. gutturalis (p. 17), S. minima (p. 22), S. minutissima (p. 29), and S. punctiventris (p. 34).Philip A. Silverstone-Sopkin (Universidad del Valle, Sección de Botánica) generously provided crucial locality and other data for the material he collected in the late 1960s and early 1970s. Robert Bezy and David Kizirian (LACM), John E. Cadle (MCZ), and William E. Duellman and Linda Trueb (KU) made sizeable loans of specimens under their care. In Colombia, Santiago Ayerbe (MHNUC), Fernando Castro (UVC), John D. Lynch (ICN), Yaneth Muñoz (IAvH), and Vivian Páez (MHNUA) provided space and access to specimens, and John D. Lynch provided long-distance confirmations of several observations. Lucas Barrientos (MHNUA and ICN) went to great lengths to collect specimens of Silverstoneia erasmios to be included in this study. John E. Cadle (MCZ) and John Simmons (KU) provided important information in our attempt to track down the holotype of Silverstoneia flotator. We are especially grateful to Kent Beaman and Jeff Seigel (LACM) for their patience in answering our many requests for additional data.  Grant's field work with Silverstoneia was facilitated by Wilmar Bolívar-G., David Fajardo, David Huff, Pablo Lehmann, Juán Pablo López, Jaime Ramírez, and Martha Solís. The Boroboro Emberó community, led by Efrén Chamorro, willingly shared their knowledge of local fauna and granted access to their territory. Grant's initial work on this project was funded by an American Museum of Natural History Collection Study Grant in 1998 and subsequently by Graduate Student Fellowships from the American Museum and the Center for Environmental Research and Conservation (CERC), Columbia University, an NSF Doctoral Dissertation Improvement Grant (DEB 0309226; awarded to T.G. and D.R. Frost), Conselho Nacional de Desenvolvimento Científico e Tecnológico Proc. 307001/2011-3, and Fundação de Amparo à Pesquisa do Estado de São Paulo Proc. 2012/10000-5. Fieldwork in the Chocó was partially funded through donations to the Kelowna Museum by Larry Bell and the Bond family; funding was organized at the Kelowna Museum by Dan Bruce. Equipment was provided by a grant from Idea Wild. Finally, special thanks are reserved for Fernando Castro (UVC), who provided direct support for Grant's research and whose efforts to facilitate and carry out herpetological research in western Colombia continue to increase our understanding of the diversity of amphibians and reptiles in that crucially important region.  Myers's fieldwork in Chocoan Colombia was initiated in 1970 in collaboration with the late John W. Daly—under the auspices of the American Museum of Natural History, the National Institutes of Health, and the late F. Carlos Lehmann, then director of the Museo Departamental de Historia Natural in Calí.  Sources of financial support for the Myers-Daly Colombian expeditions included contracts and grants to Myers from the National Institutes of Health, the Lincoln Ellsworth Fund of the American Museum of Natural History, the Camille and Henry Dreyfus Foundation, the Merck Sharp & Dohme Research Laboratories, Astra Läkemedal AB (Sweden), and the Phipps Foundation. Myers also is grateful for courtesies rendered in support of this work by authorities in the Seccion de Recursos Naturales, CVC (Corporacion Autonoma Regional del Cauca). Euclides Posada Montealegre provided valuable field assistance over the years.  This study was completed with support from Robert G. Goelet, former president and Chairman Emeritus of the Board of Trustees of the American Museum of Natural History, whose funding allowed the preparation of illustrations. Pen-and-ink drawings were executed by Patricia J. Wynne; photographs of most preserved specimens were taken by Peter Goldberg. For reading and commenting on the manuscript, we are grateful to W. Ronald Heyer, William M. Lamar, and Jay M. Savage

Review of the Frog GenusSilverstoneia, with Descriptions of Five New Species from the Colombian Chocó (Dendrobatidae: Colostethinae)