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members of the family Histeridae are ecologically diverse, with body forms that are well tailored to their preferred habitats. Among these, none exhibit more strikingly specialized morphological features than those that live in the colonies of social insects, the socalled myrmecophiles and termitophiles. While numerous lineages of Histeridae have become adapted for this lifestyle, two of these stand out in particular: the Hetaeriinae and Chlamydopsinae, together comprising over 400 described species (Mazur 1997). Neither subfamily has been particularly well-studied, but the former was the subject of a recent generic revision (Helava et al. 1985). The Chlamydopsinae, however, remain very poorly known. The Chlamydopsinae is a highly distinctive, undoubtedly monophyletic group of histerids. The primary synapomorphy for the lineage (from which the subfamily name is derived) is the form of the antennal scapes: they are inserted high on the frons and form shield-like plates that cover the eyes when the head is retracted. In addition, most species exhibit remarkable excretory structures on the elytral humeri, referred to in the chlamydopsine literature as ‘epaulettes,’ due to their position. These structures, more generally known as trichomes, are the hallmarks of myrmecophilous beetles, and probably serve to disseminate appeasement or recognition substances (Seyfried 1928, Rettenmeyer 1961, Akre 1968, Kistner 1982, Holldobler & Wilson 1990), although it should be emphasized that the actual interactions of myrmecophiles with their hosts have been studied in few beetles. Most species of Chlamydopsinae have been described from Australia (Mazur 1997), and the group is clearly centered on the Australian continent, with additional species known from Tasmania, New Guinea, Fiji, India, and Japan. Thus major discontinuities exist in the known distribution of the group. Collections made by the staff of The Natural History Museum during the ‘Project Wallace’ expedition to Sulawesi in 1985 contained numerous Chlamydopsinae representing the first known records from Indonesia. The species described herein, in addition to several being described separately from Borneo (Degallier & Caterino in prep.), therefore fill an important gap in the knowledge of Chlamydopsinae. This lineage also represents one of few known Australiancentered groups to have apparently spread across Wallace’s Line, and into southeast Asia (Knight & Holloway 1990 and references therein). The higher level classification of the Chlamydopsinae has not been assessed since the works of Lea

Descriptions of the First Chlamydopsinae (Coleoptera: Histeridae) from Wallacea