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The Carboniferous – early Permian plant genus Lesleya is a characteristic component of “dryland floras” that occupied a wide range of moisture-stressed, well-drained environments in tropical regions of Euramerica. Fossil records of Lesleya are almost exclusively found in basinal lowlands. For example, occurrences in Early Pennsylvanian-age, seasonally dry, parautochthonous deposits in basinal lowlands of North America (e.g., Illinois Basin, USA) indicate that Lesleya lived in that region in low-altitude (lowland) paleoenvironments during dry climatic intervals. In this paper, we document the first occurrence of Lesleya during the Carboniferous on the Iberian Massif, in lower Gzhelian (Upper Pennsylvanian) strata of the Douro Carboniferous Basin, in northwestern Portugal. This newly discovered occurrence includes a new species, Lesleya iberiensis sp. nov., recognized on the basis of natural molds of leaves. The Portuguese Lesleya fossils are from upland intramontane deposits and occur between coal beds. Th...

LesleyaLesquereux from the Pennsylvanian of the Iberian Massif: part of a dryland megaflora from the Variscan orogen, northwestern Portugal