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The specification of the reducing conditions which determine the growth of obligate anaerobes has been the object of many investigations. Early observers stressed the maximum oxygen pressure under which the organisms would grow (Clark, 1924; McLeod, 1931) as well as the enhancement of growth by the addition of reducing substances to the medium (Quastel and Stephenson, 1926; Wurmser, 1930). Then, after the theory of reversible oxidation-reduction potentials had been developed (Cannan, Cohen, and Clark, 1926), several attempts were made to determine in the medium a limiting potential value above which anaerobes were unable to multiply. Aubel, Aubertin, and Genevois (1929), working with eight different spore-forming anaerobes and measuring potentials with dyes, concluded that an rH above twelve (Eh of-0.060 v. at pH 7) was incompatible with anaerobic growth. Plotz and Geloso (1930) working with electrodes and dyes, established for Clostridium tetani a maximum oxidationreduction potential of +0.036 v. at pH 7.0 (rH of 15). Knight and Fildes (1930) poising their potentials with a N2, 02 mixture, found positive limit of +0.110 v. at pH 7.2 (rH of 17) for Clostridium tetani. Knaysi and Dutky (1936) worked with a butanolproducing Clostridium and attempted to separate the two

The Oxidation-Reduction Potential Requirements of a Non-Spore-Forming, Obligate Anaerobe