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The order of gene loci in thephoA-phoR region of theEscherichia coli K-12 linkage map was demonstrated to belac phoA proC phoR . The end of thephoA locus corresponding to the amino terminus of alkaline phosphatase was shown to be the end nearerproC . Translation (and transcription) ofphoA is therefore in the anticlockwise direction relative to the conventionalE. coli linkage map.

Location and Orientation of the phoA Locus on the Escherichia coli K-12 Linkage Map