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The investigation reported in this paper had the two-fold objective of determining (a) if any relationship existed between salt accumulation in the barley plant and the hourly march or levels of carbohydrate accumulation ; and (b) the relation between cyclic changes in transpiration rates and the accumulation of the chemical constituents of the substrate in the shoots of barley plants. Measurements were made of the concentrations of sugars, starch, and inorganic constituents in 5-week-old barley plants at 4-hour intervals over a 24-hour cycle. The three sets of plants compared were grown in sand cul- tures supplied with solutions as follows ; (a) base nutrient only ; (b) base nutrient plus 100 milliequivalents per liter of added chloride ; and (c) base nutrient plus 200 milliequivalents per liter of added sulphate. Materials and methods Barley was used as the experimental plant ; seed of the Sacramento vari- ety was supplied by the University of California. The plants were grown at Riverside, California, in the large, out-of-door, automatically-operated sand culture equipment described by EATON (7). The seeds were planted on April 2,1940, at a depth of one inch, spaced one inch apart in the 30-inch rows, and thinned on April 23 to fifteen plants per row. The sixteen rows of barley per bed were spaced 10.5 inches apart with end rows set 5.5 inches from the ends of the beds. The composition of the three culture solutions is shown in table I. The nutrient solutions were automatically pumped onto the beds twice 1 Contribution no. 20 of the U. S. Regional Salinity Laboratory, Bureau of Plant

EFFECT OF SALINE SUBSTRATE ON HOURLY LEVELS OF CARBOHYDRATES AND INORGANIC CONSTITUENTS OF BARLEY PLANTS