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We argue that the paradoxal softness of the red blood cells (RBC) influctuation spectra experiments is apparent. We show that the effective surfaceshear modulus $\mu_s$ of the RBC obtained from fluctuation data and thatmeasured in static deformation experiments have the same order of magnitude. Asimple micromechanical model of the RBC developped for this purpose accountsfor the influence of a finite-thickness cytoskeleton on the fluctuations of thecomposite membrane-cytoskeleton system. The spectrin network cytoskeleton withthe bulk shear modulus estimated as $\mu\approx105\div 165$ Pa contributes toboth normal and tangent fluctuations of the system and confines thefluctuations of the lipid membrane. The ratio of mean square amplitudes of theRBC normal and tangent fluctuations $<X_n^2> /< X_t^2>$ calculated in the frameof the model is 2-3 orders of magnitude smaller that it is in the free membranewith the same bending and shear moduliComment: 14 pages, 4 figure

Cytoskeleton Influence on Normal and Tangent Fluctuation Modes in the Red Blood Cells