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SUMMARY A particle weight of 8.5 ° × I@ for encephalomyocarditis virus was derived by combining results from (a), sedimentation velocity measurements on highly purified preparations of virus (o-I to 5 mg./ml.) in o.I M-KCI+o-o2 M-phosphate buffer, pH 8.o, which gave an S°0,~. of i62. 3 s, (b), differential sedimentation studies of virus in the same solvent but containing various ratios of D20 to H20 which gave a partial specific volume of o'678 ml./g., with results from (e), analysis of boundary spreading during low speed centrifugation of dilute virus solutions which led to a diffusion coefficient D20,~- of I'44 × Io -7 cm.2/sec. Sedimentation equilibrium studies on similar virus preparations in the same solvent gave a particle weight of 8-52 x I o ~ . This suggested a hydrated particle diameter for the virus of 29"8 nm. and a frictional ratio fifo of I. 13o which was consistent with that of a hydrated sphere containing 0.29 g. water/g, dry virus.

Particle Weight and other Biophysical Properties of Encephalomyocarditis Virus