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It is currently held that the sporophyte, or neutral generation in archegoniate plants, is the result of elaboration of the zygote : that while in certain Algæ the zygote simply divides to form a number of spores (carpospores), in the lower Bryophyta there has been a differ­entiation of an external, sterile, and protective wall, distinct from the internal mass of spores. The higher Bryophyta show a larger proportion of sterile tissue, which in them composes the seta, columella, and wall, while the sporogenous tissue is comparatively reduced in bulk, though still forming one united band.

II. Studies in the morphology of spore-producing members. Preliminary statement on the lycopodinæ and ophioglossaceæ