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Leaf mass per unit area (LMA) and internal leaf anatomy often affect net gas exchange because of their effects on internal CO 2  conductance to the site of carboxylation, internal shading, competition for CO 2  among carboxylation sites, nitrogen concentration and its partitioning. To evaluate effects of LMA and leaf anatomy on CO 2  assimilation, water-use efficiency (WUE) and nitrogen-use efficiency (NUE), we measured LMA, leaf thickness, the thickness of mesophyll components, and gas exchange rates at ambient CO 2  concentration in leaves of six woody deciduous and evergreen species with different leaf life spans. In two species, CO 2  assimilation was also estimated at saturating CO 2  concentrations.

Internal leaf anatomy and photosynthetic resource-use efficiency: interspecific and intraspecific comparisons