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Beech ( Fagus sylvatica  L.) seedlings were cultivated from seeds sown in pots or directly in the ground in outdoor chambers that were transparent to solar radiation, and provided either ambient air or CO 2 -enriched air (ambient + 350 μmol mol −1 ). The rooting volume was high in all experiments. In the short-term experiment, potted plants were assigned to a factorial CO 2  × nutrient treatment (optimal nutrient supply and severe nutrient shortage) for 1 year. In the long-term experiment, plants were grown directly in the ground and received an optimal supply of water and nutrients in both CO 2  treatments for 3 years.

Leaf photosynthetic characteristics of beech ( Fagus sylvatica ) saplings during three years of exposure to elevated CO 2 concentration