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First rotation  Pinus radiata  D. Don trees were grown on a nitrogen-deficient sand dune in an experimental design that included lupin ( Lupinus arboreus  Sims) as an understory species, and biennial application of balanced fertilizer in a replicated split-plot factorial design with thinning treatments as subplots. From an initial 2222 stems ha −1 , stands were thinned to 1483 and 741 trees ha −1  at ages 2 and 4 years, respectively. Mean annual production of foliage, branches, stem, coarse roots, and cones was estimated from measurements of the dry matter content at ages 7, 10, 11 and 17 years. A model was used to estimate total above- and belowground dry matter production from measurements of leaf area index and foliar nitrogen concentration. An estimate of fine root production was obtained from the difference between the model estimate of total production and the estimate of aboveground production.

Carbon partitioning in Pinus radiata stands in relation to foliage nitrogen status