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probably due to supercoiling phenomena of the DNA doublehelix.We conclude that for samples critical for DNA extraction,drowning must not be used as a method of relaxation and thatcooling the samples during drowning does not improve the con-ditions. For studies that require material for both histological/dissection and molecular purposes, we suggest that a tiny pieceof tissue is cut from the animals prior to the drowning andplaced in 10–20 vols of alcohol or is frozen. For molecular studies only very little tissue is usually needed (1–3 mm

journal of molluscan studies

Microscale genetic differentiation along the vertical shore gradient in White Sea snails Littorina saxatilis (Olivi) assessed by microsatellite markers