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1Saitama Study Center, The Open University of Japan, Ômiya-ku 330–0853, Japan 2Entomology Laboratory, Zoology Division, Research Center for Biology, Indonesian Institute of Sciences (LIPI), Cibinong 16911, Indonesia 3Institute for Agro-Environmental Sciences, NARO, Tsukuba, Ibaraki 305-8604, Japan 4Graduate School of Bioagricultural Sciences, Nagoya University, Chikusa-ku, Nagoya 464–8601 Japan 5Department of Agricultural and Environmental Biology, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Yayoi, Bunkyo-ku, Tokyo 113–8657, Japan

Phylogeography of the Delias hyparete species group (Lepidoptera: Pieridae): complex historical dispersals into and out of Wallacea