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Summary: The ModuLand plug-in provides Cytoscape users an algorithm fordetermining extensively overlapping network modules. Moreover, it identifiesseveral hierarchical layers of modules, where meta-nodes of the higherhierarchical layer represent modules of the lower layer. The tool assignsmodule cores, which predict the function of the whole module, and determineskey nodes bridging two or multiple modules. The plug-in has a detailedJAVA-based graphical interface with various colouring options. The ModuLandtool can run on Windows, Linux, or Mac OS. We demonstrate its use on proteinstructure and metabolic networks. Availability: The plug-in and its user guidecan be downloaded freely from: http://www.linkgroup.hu/modules.php. Contact:csermely.peter@med.semmelweis-univ.hu Supplementary information: Supplementaryinformation is available at Bioinformatics online.

ModuLand plug-in for Cytoscape: determination of hierarchical layers of overlapping network modules and community centrality