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The 1.69 ms spin period of PSR J1227-4853 was recently discovered in radioobservations of the low-mass X-ray binary XSS J12270-4859 following theannouncement of a possible transition to a rotation-powered millisecond pulsarstate, inferred from decreases in optical, X-ray, and gamma-ray flux from thesource. We report the detection of significant (5$\sigma$) gamma-ray pulsationsafter the transition, at the known spin period, using ~1 year of data from theLarge Area Telescope on board the Fermi Gamma-ray Space Telescope. Thegamma-ray light curve of PSR J1227-4853 can be fit by one broad peak, whichoccurs at nearly the same phase as the main peak in the 1.4 GHz radio profile.The partial alignment of light-curve peaks in different wavebands suggests thatat least some of the radio emission may originate at high altitude in thepulsar magnetosphere, in extended regions co-located with the gamma-rayemission site. We folded the LAT data at the orbital period, both pre- andpost-transition, but find no evidence for significant modulation of thegamma-ray flux. Analysis of the gamma-ray flux over the mission suggests anapproximate transition time of 2012 November 30. Continued study of the pulsedemission and monitoring of PSR J1227-4853, and other known redback systems, forsubsequent flux changes will increase our knowledge of the pulsar emissionmechanism and transitioning systems.

Discovery of Gamma-ray Pulsations from the Transitional Redback PSR J1227-4853