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An attempt was made at Clark University in 1916 and 1917 to test experimentally the thesis that the rate of physiological conduction in irritable tissues is a direct function of the electrical conductivity of the medium in which the tissue acts. The investigation was a series of experiments with reference to the propagationvelocity of the contraction-wave in the heart muscle of the river terrapin under artificial conditions. Measurements of the velocity of the wave in a series of isotonic balanced salt solutions of graded activity (mixtures of Ringer's solution with isotonic sugar solution) were made by means of special apparatus. These preliminary results indicated a direct correlation between the propagation-velocity and the electrical conductivity, and that the ratio of velocity with respect to conductivity was nearly a constant until the concentration of the salts of the Ringer's solution was reduced more than half. The original apparatus has been modified and the investigation has been extended during 1919 and 1921 to include the heart of the king-crab, Limulus, and the, sartorius of the leopard-frog, Rana pipiens. Observations have been made in the above solutions over an extended range of electrical conductivity and under nearly constant conditions of oxygen tension, hydrogen ion concentration, and temperature, t Comparative observations upon the spreading of an excitation state in irritable tissues indicate that the speed of conduction is

CORRELATION OF THE PROPAGATION-VELOCITY OF THE CONTRACTION-WAVE IN MUSCLE WITH THE ELECTRICAL CONDUCTIVITY OF THE SURROUNDING MEDIUM