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Theoretical models developed to explain the coupled solute-water transport in various absorptive epithelia consider the movement of water to be a passive process produced by the localized secretion of ions into channels associated with the epithelium (cf. 6, 9-13, 20, 28) . This standing gradient hypothesis has been extended to Malpighian tubules of insects (7, 14) . In these particular cells, osmotic gradients are thought to be established within extracellular compartments associated with basal infoldings of the plasma membrane as well as between microvilli on the lumenal surface of these cells (7, 11-14, 17) . Essential to the validity of the standing gradient hypothesis is that high molecular weight substances such as blood proteins be prevented from accumulating within

THE PERMEABILITY OF DRAGONFLY MALPIGHIAN TUBULE CELLS TO PROTEIN USING HORSERADISH PEROXIDASE AS A TRACER