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Phosphatidylinositol 4,5-bisphosphate (PI(4,5)P 2 ) is a negatively charged phospholipid that plays a major role in recruiting and regulating proteins at the plasma membrane-cytosol interface. In this issue, Chen et al. (2017. J. Cell Biol. https://doi.org/10.1083/jcb.201606047) demonstrate that RAS association domain family 4 (RASSF4) positively influences PI(4,5)P 2 synthesis through ARF6-dependent regulation of PIP5K.

RASSF4: Regulator of plasma membrane PI(4,5)P2