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Discrete, version 1.01b (Pagel, 1997), is a DOS-based program that can be used to estimate individual ancestral values. Discrete was written to test for correlated evolution between twodiscrete characters, and producing ancestor reconstructions in such away requires several steps. First, Discrete deals with two characters at a time; therefore, for thepurposes of estimating ancestors for a single character, the states must be entered twice. For example, for a tip (tipA) in state 1 on a branch 2units long andwhose immediate ancestor is nodeA, the entry would be “tipA, nodeA, 2, 1,1.” Because two characters are input, Discrete estimates two (or four) rates: a 1 and b 1 refer to q01 and q10 for the rst character, and a 2 and b 2 refer to q01 and q10 for the second character. With this in mind:

Inferring Rates of Change in Flower Symmetry in Asterid Angiosperms