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We studied a Middle Pleistocene molluscan association from northern Tenerife. It was composed of 14 species, all found in the TL-1 and TL-2 stratigraphic beds of the middle Pleistocene Unit I. The families present were Helicidae, Enidae with two new species, Pomatiidae, Canariellidae, Vitrinidae and Geomitridae. For the Geometric Morphometrics (GM) analysis, a total of 58 pictures from three localities were analysed, comparing the morphological similarities between  Napaeus guanche  sp. nov., and some closely related extant species,  Pomatias laevigatus  and extant and fossil  Hemicycla eurythyra  based on shell shape. According to the PCA,  Napaeus  accounted for 77.53% of the variation of the shell shape within the first two components,  Hemicycla  accounted for 62.62% and  Pomatias  accounted for 52.79%. The permutational ANOVA showed to be a useful analysis to discriminate between groups, presenting non-significance for  Napaeus , and significance for  Hemicycla  and  Pomatias  depicting high similarity. Despite this, the dispersion analysis showed a relatively high dispersion of the dataset for  Napaeus  and  Pomatias , related to sample size, while for  Hemicycla , the result was significant and congruent with permutational ANOVA. We describe two new species,  Napaeus lipauges  and  Napaeus guanche  increasing the known diversity of the genus  Napaeu s in the Canary Islands.

historical biology

Review of the Middle Pleistocene molluscan association from La Mancha de la Laja, Tenerife, Spain, with the description of two new species of  Napaeus  Albers, 1850 (Gastropoda: Enidae)