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From Middle-Late Triassic continental deposits of the Timezgadiouine Formation (Argana Basin, Morocco), 17 lithofacies are identified, with alluvial fan, fluvial, floodplain and lacustrine associations. Invertebrate traces include  Archaeonassa fossulata, Arenicolites  isp., cf.  Arenituba, Beaconites  isp., cf.  Camborygma, Cruziana problematica, Diplocraterion  isp.,  Fuersichnus  isp.,  Helminthoidichnites tenuis, Lockeia  isp.,  Palaeophycus striatus, Palaeophycus tubularis, Planolites  isp.,  Rusophycus carbonarius, Scoyenia gracilis, Skolithos  isp.,  Spongeliomorpha carlsbergi, Taenidium  isp. and  Taenidium barretti . Additionally, rhizoliths were observed. Six ichnoassemblages were distinguished: overbank facies with (1) Rhizolith ichnoassemblage in a low-energy floodplain environment and (2)  Scoyenia-Palaeophycus  ichnoassemblage in crevasse splay deposits. The fluvial ichnoassociation is of moderate ichnodiversity consisting of the following assemblages: (3)  Arenicolites , developed under higher flow velocities of active channels, (4)  Taenidium-Scoyenia , formed by the colonisation of inactive channels, (5)  Arenicolites -horizontal meniscate burrows, reflecting a rapid transition between active and abandoned channels and (6)  Lockeia -vertebrate footprint, formed in littoral lakes. The Aglegal and Irohalene members indicate climatic shifts between arid dry, semiarid and subhumid in a dryland river environment. The sudden onset and the high frequency of lacustrine deposits in the lower Irohalene Member point to a rapid climate change from dry to intermittent wet and dry. This change is discussed in terms of the Carnian Pluvial Event.

Ichnodiversity and facies of Triassic red beds in the Irohalene area (Argana Basin, Western High Atlas, Morocco): implications for palaeoenvironment