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At the famous Moscovian locality of Piesberg quarry near Osnabrück (Lower Saxony, Germany) there is rich abundance of Palaeodictyopterida comprising not only adults but also larval stages and their exuviae. The representatives of three palaeodictyopterid orders, eight families and 13 species were previously identified from this locality. In this study, we describe a new megascopteran  Piesbergptera niesseae  gen. nov. et sp. nov. (Piesbergpteridae fam. nov.) with some venation traits shared with Foririidae and Ischnoptilidae, both of which are known from Commentry, and new palaeodictyopterans  Osnabrugapteron rasnitsyni  gen. nov. et sp. nov. (Archaemegaptilidae),  Spilaptera peckae  sp. nov. (Spilapteridae) and  Saxonyptilus sowiaki  gen. nov. et sp. nov. (Spilapteridae), the latter providing the first evidence of nygmata among spilapterids. Based on the evaluation of forewing venation, we propose the restoration of the genus  Dictyoneurella  as a member of the palaeodictyopteran family Archaemegaptilidae.

historical biology

New palaeodictyopterid insects from the Pennsylvanian of Piesberg reveal spectacular disparity in the patterns of the wing venation (Megasecoptera, Palaeodictyoptera)